intricate shells of calcium carbonate – corals, algae, and shellfish record the conditions of the water in which they live as they grow. The chemistry of these structures carries information about the environment [...] example is the isotopic composition of boron, which reflects the pH of ancient seawater – a measure of ocean acidity that is closely linked to atmospheric CO₂ levels. These are just two of the signals we [...] conditions, we need to understand the physiological environment in which the shell forms: how calcium and other ions reach the site of calcification, how the organism regulates the chemistry at that site for